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Since their emergence over 200,000 years ago, modern humans have established homes and communities all over the planet.
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But they didn't do it alone.
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Whatever corner of the globe you find homo sapiens in today, you're likely to find another species nearby: canis lupus familiaris.
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Whether they're herding, hunting, sledding, or slouching, the sheer variety of domestic dogs is staggering.
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But what makes the story of man's best friend so surprising is that they all evolved from a creature often seen as one of our oldest rivals: canis lupus, or the gray wolf.
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When our Paleolithic ancestors first settled Eurasia roughly 100,000 years ago, wolves were one of their main rivals at the top of the food chain.
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Able to exert over 300 lbs. of pressure in one bone-crushing bite and sniff out prey more than a mile away, these formidable predators didn't have much competition.
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Much like human hunter-gatherers, they lived and hunted in complex social groups consisting of a few nuclear families, and used their social skills to cooperatively take down larger creatures.
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Using these group tactics, they operated as effective persistence hunters, relying not on outrunning their prey, but pursuing it to the point of exhaustion.
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But when pitted against the similar strengths of their invasive new neighbors, wolves found themselves at a crossroads.
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For most packs, these burgeoning bipeds represented a serious threat to their territory.
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But for some wolves, especially those without a pack, human camps offered new opportunities.
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Wolves that showed less aggression towards humans could come closer to their encampments, feeding on leftovers.
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And as these more docile scavengers outlasted their aggressive brethren, their genetic traits were passed on, gradually breeding tamer wolves in areas near human populations.
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Over time humans found a multitude of uses for these docile wolves.
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They helped to track and hunt prey, and might have served as sentinels to guard camps and warn of approaching enemies.
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Their similar social structure made it easy to integrate with human families and learn to understand their commands.
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Eventually they moved from the fringes of our communities into our homes, becoming humanity's first domesticated animal.
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The earliest of these Proto-Dogs or Wolf-Dogs, seem to have appeared around 33,000 years ago, and would not have looked all that different from their wild cousins.
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They were primarily distinguished by their smaller size and a shorter snout full of comparatively smaller teeth.
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But as human cultures and occupations became more diverse and specialized, so did our friends: short stocky dogs to herd livestock by nipping their heels; elongated dogs to flush badgers and foxes out of burrows; thin and sleek dogs for racing; and large, muscular dogs for guard duty.
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With the emergence of kennel clubs and dog shows during England's Victorian era, these dog types were standardized into breeds, with many new ones bred purely for appearance.
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Sadly, while all dog breeds are the product of artificial selection, some are healthier than others.
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Many of these aesthetic characteristics come with congenital health problems, such as difficulty breathing or being prone to spinal injuries.
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Humanity's longest experiment in controlled evolution has had other side effects as well.
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Generations of selection for tameness have favored more juvenile and submissive traits that were pleasing to humans.
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This phenomenon of selecting traits associated with youth is known as neoteny, and can be seen in many domestic animals.
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Thousands of years of co-evolution may even have bonded us chemically.
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Not only can canines understand our emotions and body language, but when dogs and humans interact, both our bodies release oxytocin, a hormone commonly associated with feelings of love and protectiveness.
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It might be difficult to fathom how every Pomeranian, Chihuahua, and Poodle are descended from fierce wolves.
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But the diversity of breeds today is the result of a relationship that precedes cities, agriculture, and even the disappearance of our Neanderthal cousins.
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And it's heartening to know that given enough time, even our most dangerous rivals can become our fiercest friends.